Answer :

In an ecosystem, the biotope refers to the physical environment where living organisms inhabit. When comparing the biotopes of the upper and lower courses of a river, there are several key differences:

1. Water Flow:
- In the upper course, the water flow tends to be faster and more turbulent due to the steep terrain and higher elevation, leading to erosion and a rocky substrate.
- In the lower course, the water flow is slower and more stable, with a wider channel and sediment deposition forming sandy or muddy substrates.

2. Temperature:
- The upper course generally experiences cooler temperatures as it is closer to the water source and may be shaded by surrounding vegetation.
- The lower course tends to have warmer temperatures due to receiving more sunlight and being influenced by wider climatic conditions.

3. Nutrient Levels:
- The upper course typically has lower nutrient levels as there is less organic matter and nutrient input from upstream sources.
- The lower course tends to have higher nutrient levels due to the accumulation of organic matter from tributaries and human activities.

4. Biodiversity:
- The upper course may have species adapted to fast-flowing water and rocky habitats, such as certain species of fish and invertebrates.
- The lower course can support a wider variety of species including fish, plants, and birds adapted to slower water flow and different substrate types.

5. Human Impact:
- Human activities like agriculture, urbanization, and pollution tend to be more pronounced in the lower course of rivers, impacting water quality and biodiversity.
- The upper course is often less affected by human intervention, maintaining a more pristine environment in some cases.

Understanding these differences in the biotopes of the upper and lower courses of a river is crucial for conservation efforts and ecosystem management to ensure the health and sustainability of river ecosystems.